Fish macro- and microanatmy, in particular the structure and development of the skeletal system.

BIO 280 Fish biology: Anatomy part

BIO 381 Fish histopathology: Normal fish histology

(BIO 270 Vertebrate anatomy: Macroanatomy

BIO 305 Methods in microscopy)

Lecture

• Harald Kryvi (1996). Anathomy and phylogeny of the carps. (external link)
• Marius Takvam; Kristina Sundell; Henrik Sundh et al. (2023). Modifikasjoner av Na+, K+ - ATPase enzymaktivitets protokollen og bruksområder i moderne lakseoppdrett. (external link)

Academic article

• Harald Kryvi; A. Aarsland; Rolf Kristian Berge (1990). Morphologic effects of sulfursubstituted fatty acids on rat hepatocytes with special references to proliferation of peroxisomes and mitochondria. (external link)
• Geir Kåre Totland; Lise Madsen; Beate Klementsen et al. (2000). Proliferation of mitochondria and gene expression of carnitine palmitoyltransferase and fatty acyl-CoA oxidase in rat skeletal muscle, heart and liver by hypolipidemic fatty acids. (external link)
• Geir Kåre Totland; Per Gunnar Fjelldal; Harald Kryvi et al. (2011). Sustained swimming increases the mineral content and osteocyte density of salmon vertebral bone. (external link)
• G.O. Taranger; C. Haux; S.O. Stefansson et al. (1999). Mechanisms underlying photoperiodic effects on age at sexual maturity in Atlantic salmon, Salmo salar L. (external link)
• Per Gunnar Fjelldal; Geir Kåre Totland; Tom Johnny Hansen et al. (2013). Regional changes in vertebra morphology during ontogeny reflect the life history of Atlantic cod (Gadus morhua L.). (external link)
• Livar Frøyland; Knut Helland; Harald Kryvi et al. (1996). Increased fatty acid oxidation and polydispersity of rat liver mitochondria and peroxisomes induced by the hypolipidemic agent tetradecylthioacetic acid. (external link)
• Sindre Grotmol; Geir Kåre Totland; Harald Kryvi et al. (2002). Spatial distribution of muscle fibre types within skeletal muscle fascicules from standardbred horses. (external link)
• Marta Eide; Odd Andre Karlsen; Harald Kryvi et al. (2014). Precision-cut liver slices of Atlantic cod (Gadus morhua): An in vitro system for studying the effects of environmental contaminants. (external link)
• Shou Wang; Harald Kryvi; Sindre Grotmol et al. (2013). Mineralization of the vertebral bodies in Atlantic salmon (Salmo salar L.) is initiated segmentally in the form of hydroxyapatite crystal accretions in the notochord sheath. (external link)
• Sindre Grotmol; Harald Kryvi; Kari Nordvik et al. (2003). Notochord segmentation may lay down the pathway for the development of the vertebral bodies in the Atlantic salmon. (external link)
• Per Gunnar Fjelldal; Sankar Murugesan; Tone Vågseth et al. (2025). Vertebral deformities in cultured big size Rainbow Trout: Radiological analysis from juvenile to harvest size. (external link)
• Odd Børre Humborstad; Keno Ferter; Harald Kryvi et al. (2016). Exophthalmia in wild-caught cod (Gadus morhua L.): development of a secondary barotrauma effect in captivity. (external link)
• Harald Kryvi; Iselin Rusten; Per Gunnar Fjelldal et al. (2017). The notochord in Atlantic salmon (Salmo salar L.) undergoes profound morphological and mechanical changes during development. (external link)
• Sindre Grotmol; Geir Kåre Totland; Harald Kryvi (2005). Deformation of the notochord by pressure from the swim bladder may cause malformation of the spine in reared Atlantic cod larvae: a case study. (external link)
• Kari Nordvik; Harald Kryvi; Geir Kåre Totland et al. (2005). The salmon vertebral body develops through mineralization of two preformed tissues that are encompassed by two layers of bone. (external link)
• Sindre Grotmol; Kari Nordvik; Harald Kryvi et al. (2005). A segmental pattern of alkaline phosphatase activity with the notochord coincides with the initial formation of the vertebrae. (external link)
• Erik Slinde; Pernille Baardseth; Harald Kryvi (1983). Effect of ascorbat on a hypotenically isolated mitochondrial fraction during ageing of Swede (Brassica napus var. napobrassica L). (external link)
• Antony Jesu Prabhu Philip; Thomas Fraser; Kari Nordvik et al. (2024). Sequence of formation and inheritance of meristic variation in the post-cranial axial skeleton of Atlantic salmon (Salmo salar). (external link)
• Geir Lasse Taranger; C. Haux; Tom Johnny Hansen et al. (1999). Mechanisms underlying photoperiodic effects on age at sexual maturity in Atlantic salmon, Salmo salar. (external link)
• Einar Ringø; Terry Mayhew; Harald Kryvi et al. (2001). Damaging effect of dietary inulin to intestinal enterocytes in Arctic charr (Salvelinus alpinus L.). (external link)
• Olav Sigurd Kjesbu; Harald Kryvi; Birgitta Norberg (1996). Oocyte size and structure in relation to blood plasma steroid hormones in individually monitored, spawning Atlantic cod. (external link)
• Einar Ringø; Terry Mayhew; Harald Kryvi et al. (2001). Damaging effect of dietary inulin to intestinal enterocytes in Arctic charr (Salvelinus alpinus L.). (external link)
• Anita Sagstad; Sindre Grotmol; Harald Kryvi et al. (2011). Identification of vimentin- and elastin-like transcripts specifically expressed in developing notochord of Atlantic salmon (Salmo salar L.). (external link)
• Jessica Marks; Christiane H. Biermann; Walter F. Eanes et al. (2008). Sperm Polymorphism Within the Sea Urchin Strongylocentrotus droebachiensis: Divergence Between Pacific and Atlantic Oceans. (external link)
• Marius Takvam; Elsa Denker; Naouel Gharbi et al. (2021). Sulfate homeostasis in Atlantic salmon is associated with differential regulation of salmonid-specific paralogs in gill and kidney. (external link)
• Shou Wang; Tomasz Furmanek; Harald Kryvi et al. (2014). Transcriptome sequencing of Atlantic salmon (Salmo salar L.) notochord prior to development of the vertebrae provides clues to regulation of positional fate, chordoblast lineage and mineralisation. (external link)
• Per Gunnar Fjelldal; Monica Favnebøe Solberg; Tom Johnny Hansen et al. (2016). Salmonid fish: model organisms to study cardiovascular morphogenesis in conjoined twins?. (external link)
• PG Fjelldal; Sindre Grotmol; Harald Kryvi et al. (2004). Pinealectomy induces malformation of the spine and reduces the mechanical strength of the vertebrae in Atlantic salmon, Salmo salar. (external link)
• Sindre Grotmol; Harald Kryvi; Roger Keynes et al. (2006). Stepwise enforcement of the notochord and its intersection with the myoseptum: an evolutionary path leading to development of the vertebra?. (external link)
• Sankar Murugesan; Harald Kryvi; Thomas Fraser et al. (2024). A new method for regionalization of the vertebral column in salmonids based on radiographic hallmarks. (external link)
• A. Nissling; Harald Kryvi; L. Valin (1994). Variation in egg bouancy of baltic cod (Gadus morhua) and implications for egg survival at prevailing conditions in the Baltic sea. (external link)
• K. Hamre; B. Hjeltnes; Harald Kryvi et al. (1994). Decreased concentration of haemoglobin, accumulation of lipid oxidation products and unchanged skeletal muscle in Atlantic salmon (Salmo salar) fed low dietary vitamin E. (external link)
• Erik Slinde; Knut-Jan Andersen; Lars Mørkrid et al. (1976). Sedimentation Coefficient, Buoyant Density, Morphology, and Optimal Recorvery of Mitochondria from the Ileal Mucosa of Guinea Pigs. (external link)
• Sindre Grotmol; Harald Kryvi; Kari Nordvik et al. (2003). Notochord segmentation may lay the pathway for the development of the vertebral bodies of the Atlantic salmon Salmo salar. (external link)
• K. Hamre; B. Hjeltnes; Harald Kryvi et al. (1994). Decreased concentration of hemaglobin, accumulation of lipid oxidation products and unchanged skeletal muscle in atlantic salmon (Salmo salar) fed on low diatary vitamin E. (external link)
• Sankar Murugesan; Thomas Fraser; Harald Kryvi et al. (2025). Effect of heterozygosity, ploidy and incubation temperature on post-cranial axial skeletal meristics and deformities in Atlantic salmon (Salmo salar). (external link)
• Yuko Kamisaka; Ann-Elise Olderbakk Jordal; Rolf Edvardsen et al. (2010). A case report on the distended gut syndrome (DGS) in cultured larvae of Atlantic cod (Gadus morhua). (external link)
• K. Hamre; B. Hjeltnes; Harald Kryvi et al. (1994). Decreased concentration of hemoglobin, accumulation of lipid oxidation products and unchanged skeletal muscle in atlantic salmon (Salmo salar) fed on low diatary viatamin E. (external link)
• Sindre Grotmol; Kari Nordvik; Harald Kryvi et al. (2005). A segmental pattern of alkaline phosphatase activity within the notochord coincides with the initial formation of the vertebral bodies. (external link)
• Livar Frøyland; Knut Helland; Geir Kåre Totland et al. (1996). A hypolipidemic peroxisome proliferating fatty acid induces polydispersity of rat liver mitochondria. (external link)
• Harald Kryvi; Kari Nordvik; Per Gunnar Fjelldal et al. (2020). Heads and tails: The notochord develops differently in the cranium and caudal fin of Atlantic Salmon (Salmo salar, L.). (external link)
• Olav Sigurd Kjesbu; Jarle Klungsøyr; Peter R. Witthames et al. (1991). Fecundity, atresia, and egg size of captive Atlantic cod (Gadus morhua) in relation to proximate body composition. (external link)
• Sindre Grotmol; Geir Kåre Totland; Harald Kryvi et al. (2002). Spatial distribution of fiber types within skeletal muscle fascicles from Standardbred horses. (external link)
• T. F. Galloway; E. Kjørsvik; Harald Kryvi (1998). Effect of temperature and viability and axial muscle development in embryos and yolk sac larvae of the northeast arctic cod (Gadus morhua). (external link)
• Vilde Arntzen Engdal; Alf Seljenes Dalum; Harald Kryvi et al. (2024). State of the heart: Anatomical annotation and assessment of morphological cardiac variation in Atlantic salmon (Salmo salar L.). (external link)
• K. Hamre; B. Hjeltnes; Harald Kryvi et al. (1994). Decreased concentration of Hemoglobin, accumualtion of lipid oxidation products and unchanged skeletal muscle in Atlantic salmon (Salmo salar) fed low dietary vitamin E. (external link)

Conference abstract

• M E Porter; Sindre Grotmol; Harald Kryvi et al. (2012). The Vertebral Column of Sharks: Functional Morphology of the Intervertebral Joint. (external link)

Academic literature review

• Marius Takvam; Chris M. Wood; Harald Kryvi et al. (2021). Ion transporters and osmoregulation in the kidney of teleost fishes as a function of salinity. (external link)
• Marius Takvam; Chris M. Wood; Harald Kryvi et al. (2023). Role of the kidneys in acid-base regulation and ammonia excretion in freshwater and seawater fish: implications for nephrocalcinosis. (external link)
• Marius Takvam; Kristina Sundell; Henrik Sundh et al. (2023). New wine in old bottles: Modification of the Na+/K+-ATPase enzyme activity assay and its application in salmonid aquaculture. (external link)

Master’s thesis

• Egil-André Drønnesund; Harald Kryvi; Jaakko Saraste (2006). Distribution of markers of early secretory compartments in pancreatic exocrine cells. (external link)
• Nina Karin Aarskog; Stein Ove Doskeland; Harald Kryvi (1992). Effects of cAMP elevating agents and toxins on differentiation and viability of cultured neuroblastoma cells.. (external link)
• Geir Kåre Totland; Harald Kryvi; Sindre Grotmol et al. (2006). Morfologiske effekter av vaksineindusert bukhinnebetennelse på bukspyttkjertelen hos Atlanterhavslaks (Salmo salar L.). (external link)

Conference lecture

• Shou Wang; Tomasz Furmanek; Harald Kryvi et al. (2013). DEEP SEQUENCING REVEALS MINERALIZING NOTOCHORD TRANSCRIPTOME, DURING AXIAL SKELETON ENFORCEMENT FROM NOTOCHORD TO VERTEBRA IN ATLANTIC SALMON (SALMO SALAR L.). (external link)
• P. R. Flood; B. Hjeltnes; Geir Kåre Totland et al. (1994). Tidlig smittespredning fra ILA-infisert fisk. (external link)
• Oddmund Søvik; A. L. B. Monsen; Rolf Kristian Berge et al. (1994). Zellweger-like syndrome: Prolonged survival in male twins during treatment with n-3 fatty acids. (external link)
• B. Hjeltnes; P. R. Flood; Geir Kåre Totland et al. (1994). Transmission of infectious salmon anemia (ISA) throug naturally secreted material. (external link)
• Geir Kåre Totland; B. Hjeltnes; P. R. Flood et al. (1994). Naturlige smitteveier for ILA. (external link)
• Erik-Jan Lock; Christel Krossøy; Rune Waagbø et al. (2008). Roles of fat soluble vitamins in bone development and mineral metabolism. (external link)
• Robin Ørnsrud; Sindre Grotmol; Harald Kryvi et al. (2004). Roles of fat soluble vitamins in bone development and mineral metabolism. (external link)
• Robin Ørnsrud; Christel Krossøy; Rune Waagbø et al. (2006). Fettløselige vitaminers rolle i beinutvikling og mineralmetabolisme. (external link)
• Shou Wang; Tomasz Furmanek; Harald Kryvi et al. (2012). The Atlantic salmon (Salmo Salar L.) notochord displays a dynamic transcriptome prior to and during vertebral segmentation. (external link)

Working paper

• G.K. Totland; Harald Kryvi (1998). VERTEBRATENES ANATOMI. Kompendium for grunnemne i biologi ved UiB. (external link)

Doctoral thesis (PhD)

• Harald Kryvi (1978). Studies on the histochemistry, ultrastructure and growth of axial muscle fibres in the sharks Etmopterus spinax and Galeus melastomus. (external link)

Conference poster

• Rune Waagbø; Olav Breck; Harald Kryvi et al. (2006). Workshop innen beindeformiteter hos laks og torsk i intensivt oppdrett. (external link)
• Kaja H Skjærven; Roderick Nigel Finn; Harald Kryvi et al. (2002). Yolk resorption in developing plaice (Pleuronectes platessa). (external link)

Academic book chapter

• Kaja H Skjærven; Roderick Nigel Finn; Harald Kryvi et al. (2003). Yolk resorption in developing plaice (Pleuronectes platessa). (external link)
• Torgeir Flatmark; Jon Kvannes; T. S. Eikhom et al. (1992). Differential responses at the peroxisomal level in hepatocytes and myocardium on feeding rats diets containing partially hydrogenated fish oil. (external link)
• Rolf Kristian Berge; Harald Kryvi; Niels Aarsæther et al. (1993). A review in peroxisomes, biology and importance in toxicology and medicine. (external link)
• Ivar Lossius; Harald Kryvi; James Lorens et al. (1993). Cloning and characterization of growth regulating genes from Atlantic salmon: A strategy for increasing growth rate by microinjection of a modified growth hormone gene into fertilized salmon eggs. (external link)
• Rolf Kristian Berge; Harald Kryvi; Niels Aarsæther et al. (1993). Proliferation of peroxisomes and mitochondria and modulation of lipid metabolizing enzymes by hypolipidaemic non-_b-oxidizable fatty acids (3-thia fatty acids). (external link)
• I. Kristensen; Harald Kryvi; S. Mortensen (1994). Sjøpattedyr. (external link)
• Harald Kryvi; S. Mortensen (1994). Fiskenes anatomi og fysiologi. (external link)
• Harald Kryvi; H. Lysebo (1995). Higher education in biology in Norway. (external link)

Non-fiction monograph

• Harald Kryvi; Geir Kåre Totland (1995). Fiskeanatomi. (external link)

Popular science article

• Harald Kryvi (1996). Kjevebevegelsen hos fisk. (external link)
• Yuko Kamisaka; Ann-Elise Olderbakk Jordal; Rolf Edvardsen et al. (2010). Tarmlidelser hos torskelarver– hvor ligger forklaringene?. (external link)
• Sindre Grotmol; Teresa Cieplinska; Harald Kryvi et al. (2003). "Nakkeknekk" hos torskeyngel. (external link)

See a complete overview of publications in NVA.

See the list for the research group.

See the list for the research group.

Cand. real. 1970.

Dr. philos. 1978.

Professor 1982.